Research Projects

Current Projects

  • 2015-2017: CGL2014-52611-C2-1-P. "Adaptaciones anatómicas de la región maxilofacial y de la dentición en los primates en relación con la dieta y el hábitat: implicaciones adaptativas y filogenéticas". 60.000 €. PI: Dr. Alejandro Pérez-Pérez.
  • 2014-2017: CGL2014-52611-C2-2-P. "Adaptaciones del sistema musculoesquelético de la extremidad anterior de los primates a diferentes tipos de locomoción: implicaciones evolutivas y funcionales". 60.000 €. PI: Dr. JM Potau.

o    Main publications:

1.    Aliaga-Martínez A, Romero A, Galbany J, Hernández-Aguilar A & Pérez-Pérez A (2017) Buccal dental microwear texture and catarrhine diets. American Journal of Physical Anthropology. doi: 10.1002/ajpa.23219.

2.    Galbany J, Imanizabayo O, Romero A, Vecellio V, Glowacka H, Cranfield MR, Bromage TG, Mudakikwa A, Stoinski TS & McFarlin SC (2016) Tooth wear and feeding ecology in mountain gorillas from Volcanoes National Park, Rwanda. American Journal of Physical Anthropology 159: 457-465.

3.    Galbany J, Tung J, Altmann J & Alberts SC (2015) Canine size in Amboseli male baboons: maturation, wear and mating success. PLoS ONE 10(5): e0126415.

4.    Gamarra B, Nova Delgado M, Romero A, Galbany J & Pérez-Pérez A (2016) Phylogenetic signal in molar dental shape of extant and fossil catarrhine primates. Journal of Human Evolution 94: 13-27.

5.    Górka K, Romero A & Pérez-Pérez A (2015) First molar size and wear within and among modern hunter-gatherers and agricultural populations. HOMO - Journal of Comparative Human Biology 66(4): 299-315.

6.    Górka K, Romero A & Pérez-Pérez A (2016) Dental-macrowear and diet of Tigara foragers from Point Hope, northern Alaska. Anthropol Anz 73:257–264.

7.    Martínez LM, Estebaranz-Sánchez F, Galbany J & Pérez-Pérez A (2016) Testing dietary hypotheses of east african hominines using buccal dental microwear data. PLoS ONE 11(11): e0165447.

8.    Martínez LM, Estebaranz-Sánchez F, Pérez-Pérez A (2016) Buccal dental microwear as an indicator of diet in modern and ancient human populations. In: Wild harvest: Plants in the hominin and pre-agrarian human worlds (Hardy K, Kubiak-Martens L, eds.). Oxbow Books, Oxford, UK.

9.    Nova Delgado M, Galbany J & Pérez-Pérez A (2016) Molar shape variability in platyrrhine primates. Journal of Human Evolution 99:79-92.

10. Nova Delgado M, Galbany J & Pérez-Pérez A (2016) Morphometric variation of extant platyrrhine molars: taxonomic implications for fossil platyrrhines. PeerJ 4:e1967.

11. Nova Delgado M, Galbany J, Górka K & Pérez-Pérez A (2015) Taxonomic implications of molar morphology variability in capuchins. International Journal of Primatology 36: 707-727.

12. Percher A, Merceron G, Nsi Akoue G, Galbany J, Romero A, Charpentier MJE (2017) Dental Microwear Textural Analysis as an analytical tool to depict individual traits and reconstruct the diet of a primate. American Journal of Physical Anthropology. DOI: 10.1002/ajpa.23337.

13. Percher A, Romero A, Galbany J, Nsi Akoue G, Pérez-Pérez A, Charpentier MJE (2017) Buccal dental-microwear and dietary ecology in a free-ranging population of mandrills (Mandrillus sphinx) from southern Gabon. PlosOne. Accepted.

14. Pérez-Pérez A, Lozano M, Romero A, Martínez LM, Galbany J, Pinilla B, Estebaranz-Sánchez F, Bermúdez de Castro JM, Carbonell E & Arsuaga JL (2017) The diet of the first europeans from Atapuerca. Scientific Reports 7: 43319.

15. Salazar-García DC, Romero A, García-Borja P, Subirà ME, Richards MP (2016) A combined dietary approach using isotope and dental buccal-microwear analysis of human remains from the neolithic, roman and medieval periods from the archaeological site of Tossal de les Basses (Alicante, Spain). Journal of Archaeological Science 6: 610-619.


Past Projects

  • SGR 2014-2016: “Biologia Evolutiva dels Humans i altres Primats”. Programa: PIGC - Projectes de recerca per potenciar els grups de recerca consolidats. AGAUR - Agència de Gestió d'Ajuts Universitaris i de Recerca. Generalitat de Catalunya. SGR2014-45. PI: Alejandro Pérez-Pérez.
  • 2012-2014: CGL2011-22999. "Medición de la tasa de cambio morfológico en relación con el cambio climático: modelos evolutivos en primates e implicaciones filogenéticas". 60.000 €. Departament de Biologia Animal, Universitat de Barcelona. PI: Dr. Alejandro Pérez-Pérez.

o    Main publications:

1.    Arias-Martorell J, Alba DM, Potau JM, Bello-Hellegouarch G, Pérez-Pérez  A (2015) Morphological affinities of the proximal humerus of Epipliopithecus vindobonensis and Pliopithecus antiquus: suspensory inferences based on a 3D geometric morphometrics approach. Journal of Human Evolution 80: 83-95.

2.    Arias-Martorell J, Potau JM, Bello-Hellegouarch G & Pérez-Pérez A (2014) Brief communication: Developmental versus functional three-dimensional geometric morphometric-based modularity of the human proximal humerus. Am J Phys Anthropol. 2014;154(3):459-65.

3.    Arias-Martorell J, Potau JM, Bello-Hellegouarch G & Pérez-Pérez A (2015) Like father, like son: assessment of the morphological affinities of A.L. 288-1 (A. afarensis), Sts 7 (A. africanus) and Omo 199-73-1718 (Australopithecus sp.) through a three-dimensional shape analysis of the shoulder joint. PLoS ONE, 10: e0117408.

4.    Arias-Martorell J, Potau JM, Bello-Hellegouarch G, Pastor JF & Pérez-Pérez A (2012) 3D geometric morphometric analysis of the proximal epiphysis of the hominoid humerus. Journal of Anatomy 221(5):394-405.

5.    Arias-Martorell J, Tallman M, Potau JM, Bello-Hellegouarch G & Pérez-Pérez A (2015) Shape analysis of the proximal humerus in orthograde and semi-orthograde primates: correlates of suspensory behavior. American Journal of Primatology 77: 1-19.

6.    Bello-Hellegouarch G, Aziz MA, Ferrero EM, Kern M, Francis N, Diogo R (2013) Pollical palmar interosseous muscle (Musculus adductor pollicis accessorius): Attachments, innervation, variations, phylogeny, and implications for human evolution and medicine. Journal of Morphology 274(3):275-293.

7.    Bello-Hellegouarch G, Potau JM, Arias-Martorell J, Pastor JF & Pérez-Pérez A (2013) Brief communication: Morphological effects of captivity: A geometric morphometric analysis of the dorsal side of the scapula in captive-bred and wild-caught hominoidea. American Journal of Physical Anthropology 152(2):306-310.

8.    Bello-Hellegouarch G, Potau JM, Arias-Martorell J, PastorJF & Pérez-Pérez A (2013) A comparison of qualitative and quantitative methodological approaches to characterizing the dorsal side of the scapula in Hominoidea and its relationship to locomotion. International  Journal of Primatology 34:315-336.

9.    Estebaranz F, Galbany J, Martínez LM, Turbón D & Pérez-Pérez A (2012) Buccal dental microwear analyses support greater specialization in consumption of hard foodstuffs for Australopithecus anamensis. Journal of Anthropological Sciences 90: 163-185.

10. Galbany J, Romero A, Mayo-Alesón M, Itsoma F, Gamarra B, Pérez-Pérez A, Willaume E, Kappeler PM & Charpentier MJE (2014) Age-related tooth wear differs between forest and savanna primates. PLoS ONE 9(4):e94938.

11. Nova Delgado M, Gamarra B, Nadal J, Mercadal O, Olesti O, Guàrdia J, Pérez-Pérez A & Galbany J (2014) Dental shape variability in cercopithecoid primates: A model for the taxonomic attribution of macaques from roman archaeological contexts. Folia Primatologica 85: 361-378.

12. Romero A, Galbany J, De Juan J & Pérez-Pérez A (2012) Short and long-term in vivo human buccal dental-microwear turnover. American Journal of Physical Anthropology 148(3): 467-472.

13. Romero A, Ramírez-Rozzi FV, De Juan J, Pérez-Pérez A (2013) Diet-related buccal dental microwear patterns in central african pygmy foragers and bantu-speaking farmer and pastoralist populations. PLoS ONE 8(12): e84804.


  • 2009-2013: Grup d’Estudis d’Evolució Dels Homínids i d’altres Primats. 54,000.00€. Programa: PIGC - Projectes de recerca per potenciar els grups de recerca consolidats. AGAUR - Agència de Gestió d'Ajuts Universitaris i de Recerca. Generalitat de Catalunya. 2009SGR884. PI: Alejandro Pérez-Pérez.
  • 2011: CGL2010-15340. "Adaptación y ambiente: modelos anatómicos y ecológicos de la dentición y el esqueleto en primates cercopitecoideos, hominoideos y poblaciones humanas cazadoras-recolectoras." 9,000.00€. Departament de Biologia Animal, Universitat de Barcelona. PI: Dr. Alejandro Pérez-Pérez.

o    Main publications:

1.    Galbany J, Altmann J, Pérez-Pérez A & Alberts SC (2011) Age and individual foraging behavior predict tooth wear in Amboseli baboons. American Journal of Physical Anthropology 144(1): 51-59.

2.    Gamarra B, Romero A, Pérez-Pérez A & Galbany J (2011) Análisis del error de medición y variabilidad interespecífica morfométrica de Fourier en M2 de primates Hominoidea. Revista Española de Antropología Física 32: 56-66.

3.    Gamarra B, Romero A, Pérez-Pérez A & Galbany J (2011) Caracterización morfológica del M2 de primates Hominoidea a partir de análisis de Fourier. Revista Argentina de Antropología Biológica 13(1): 29-41.

4.    Pinilla Pérez B, Romero A & Pérez-Pérez A (2011) Age-related variability in buccal dental-microwear in middle and upper pleistocene human populations. Anthropological Review 74: 25-37.

5.    Potau JM, Artells R, Bello G, Muñoz C, Monzó M, Pastor JF, de Paz F, Barbosa M, Diogo R & Wood B (2011) Expression of myosin heavy chain isoforms in the supraspinatus muscle of different primate species: implications for the study of the adaptation of primate shoulder muscles to different locomotor modes. International Journal of Primatology 32: 931-944.


  • 2008-2010: CGL2007-60802/BTE. "Modelos evolutivos de periodicidad temporal en Homínidos: las fluctuaciones climáticas y ecológicas y la fragmentación de hábitat como mecanismos de especiación y diversificación." 100.000,00€. Departament de Biologia Animal, Universitat de Barcelona. PI: Dr. Alejandro Pérez-Pérez.

o    Main publications:

1.    Altmann J, Gesquiere L, Galbany J, Onyango PO & Alberts SC (2010) The life history context of reproductive aging in a wild primate model. Annals of the New York Academy of Sciences 1204: 127-138.

2.    Aversi-Ferreira TA, Diogo R, Potau JM, Bello G, Pastor JF & Ashraf Aziz M (2010) Comparative anatomical study of the forearm extensor muscles of Cebus libidinosus (Rylands et al., 2000; Primates, Cebidae), modern humans, and other primates, with comments on primate evolution, phylogeny, and manipulatory behavior. The anatomical record 293: 2056-2070.

3.    Estebaranz F, Martínez LM, Galbany J, Turbón D & Pérez-Pérez A (2009) Testing hypotheses of dietary reconstruction from buccal dental microwear in Australopithecus afarensis. Journal of Human Evolution 57: 739-750.

4.    Galbany J, Dotras L, Alberts SC & Pérez-Pérez A (2010) Tooth size variation related to age in Amboseli baboons. Folia Primatologica 81(6): 348-359.

5.    Galbany J, Estebaranz F, Martínez LM & Pérez-Pérez A (2009) Buccal dental microwear variability in extant african Hominoidea primates: taxonomy versus ecology. Primates 50(3): 221-230.

6.    Pérez-Pérez A, Galbany J, Romero A, Martínez LM, Estebaranz F, Pinilla B & Gamarra B (2010) Origen y evolución de los dientes: de los cordados primitivos a los humanos modernos. Revista Española de Antropología Física 31: 167-192.

7.    Prevedorou E, Díaz-Zorita Bonilla M, Romero A, Buikstra JE, de Miguel Ibáñez MªP & Knudson KJ (2010) Residential mobility and dental decoration in early medieval Spain: Results from the eighth century site of Plaza del Castillo, Pamplona. Dental Anthropology 23 (2): 42-52.

8.    Romero A, De Miguel Ibáñez MP, Buikstra JE, Knudson KJ, Prevedorou EA, Díaz-Zorita Bonilla M & De Juan J (2009) Mutilación dentaria en la necrópolis islámica de plaza del castillo (Siglo VIII DC) de Pamplona (Navarra). Revista Española de Antropología Física 29: 1-14.

9.    Walker MJ, Gibert J, López MV, Lombardi AV, Pérez-Pérez A, Zapata J, Ortega J, Higham T, Pike A, Schwenninger JL, Zilhão J & Trinkaus E (2008) Late neandertals in southeastern Iberia: Sima de las Palomas del Cabezo Gordo, Murcia, Spain. Proceedings of the National Academy of Sciences 105(52): 20631-20636.

10. BOOK: Diogo R, Potau JM, Pastor JF, dePaz F, Ferrero EM, Bello G, Barbosa M & Wood BA (2010) Photographic and descriptive musculoskeletal atlas of gorilla: With notes on the attachments, variations, innervation, synonymy and weight of the muscles. Science Publishers.

11. BOOK: Pérez-Pérez A, Martínez LM, Estebaranz F, Galbany J & Pinilla B (2009) El llarg camí de l’evolució humana. Ed. Associació ADÉS per a la divulgació de l’Evolució Humana. Barcelona.


  • 2005-2009: Grup d’Estudis d’Evolució Dels Homínids i d’Altres Primats. Programa: PIGC - Projectes de recerca per potenciar els grups de recerca consolidats. DURSI - Departament d’Universitats, Recerca i Societat de la Informació, Generalitat de Catalunya. 2005SGR00955. 38.600,00€. PI: Adrià Casinos Pardos.
  • 2005-2006: Utilització dels Serveis de la Plataforma de Nanotecnologia del Parc Científic i dels Serveis Cientificotècnics de la Universitat de Barcelona. ACES - Accions especials de recerca. CRFB - Comisió de Recerca, Fac. Biologia UB. ACES-UB 2005. 3.000,00€. PI: Alejandro Pérez-Pérez.
  • 2005-2007: CGL2004-00775/BTE. “Análisis de las fluctuaciones de período medio y largo del microdesgaste dentario en Homínidos como indicadores paleoecológicos mediante microscopía confocal y análisis fractal sensible a escala”. 88.450,00 €. Departament de Biologia Animal, Universitat de Barcelona. PI: Dr. Alejandro Pérez-Pérez.

o    Main publications:

1.    Estebaranz F, Galbany J, Martínez LM & Pérez-Pérez A. (2007) 3-D Interferometric microscopy applied to the study of buccal enamel microwear. In: Bailey SE, Hublin JJ (Eds.) Dental perspectives on human evolution. Pp: 389-401. Springer.

2.    Galbany J, Estebaranz F, Martínez LM & Pérez-Pérez A (2007) Buccal dental microwear in extant african Hominoidea and dietary habits of Australopithecus afarensis. In: Bodzsár EB, Zsákai A (Eds.) New perspectives and problems in anthropology. Pp: 19-30. Cambridge Scholars Publishing.

3.    Galbany J, Estebaranz F, Martínez LM, Romero A, De Juan J, Turbón D & Pérez-Pérez A (2006) Comparative analysis of dental enamel polyvinylsiloxane impression and polyurethane casting methods for SEM research. Microscopy Research and Technique 69(4): 246-252.

4.    Galbany J, Martínez LM, López-Amor HM, Espurz V, Hiraldo O, Romero A, De Juan J & Pérez-Pérez A (2005) Error rates in dental buccal microwear quantification using Scanning Electron Microscopy. Scanning 27: 23-29.

5.    Galbany J, Moyà-Sola S & Pérez-Pérez A (2005) Dental microwear variability on buccal tooth enamel surfaces of extant catarrhini and miocene fossil Dryopithecus laietanus (Hominoidea). Folia Primatologica 76(6): 325-341.

6.    Gómez-Robles A, Martinón-Torres M, Bermúdez de Castro JM, Margvelashvili A, Bastir M, Arsuaga JL, Pérez-Pérez A, Estebaranz F & Martínez LM (2007) A geometric morphometric analysis of hominin upper first molar shape. Journal of Human Evolution 53(3): 272-285.

7.    Martínez LM, Pérez-Pérez A & Turbón D (2006) Nutrition in ancient hominids. In: Integrative approaches to human health and evolution. Elsevier International Congress Series 1296: 16-22.

8.    Romero & De Juan J (2007) Intra- and interpopulation human buccal tooth surface microwear analysis: inferences about diet and formation processes. Anthopologie XLV/1: 61-70.

9.    Romero A, Galbany J, Pérez-Pérez A & De Juan (2007) Microwear formation rates in human buccal tooth enamel surfaces: an experimental in vivo analysis under induced-diet. In: Bodzsár EB, Zsákai A (Eds.) New perspectives and problems in anthropology. Pp: 135-146. Cambridge Scholars Publishing.


  • 2003: “Sociedades humanas y medio natural: explotación de recursos y adaptación biológica”. Promoción Internacional de la Investigación realizada en los proyectos ACES98-7/1, ACP1999-0026 Y BMC2000-0538 para la integración en la Red de Excelencia Europea 'Cooperación and conflict resolucion; Comparative Studies on Human Social Behaviour'. Departament de Biologia Animal, Universitat de Barcelona. 9.000,00€. PI: Dr. Alejandro Pérez-Pérez.
  • 2001-2004: BMC-2000-0538. Proyectos de Investigación Científica y Desarrollo Tecnológico del Ministerio de Ciencia y Tecnología. “Modelos ecológicos de adaptación y selección en los Homínidos Plio-Pleistocénicos: la alimentación como factor determinante del crecimiento y desarrollo de las poblaciones cazadoras-recolectoras humanas”. 46.520,14€ (8.669.136 Ptes). Departament de Biologia Animal, Universitat de Barcelona. PI: Dr. Alejandro Pérez-Pérez.

o    Main publications:

 1.    Egocheaga JE, Pérez-Pérez A, Rodríguez L, Galbany J, Martínez LM & Telles Antunes M (2004) New evidence and interpretation of subvertical grooves in neandertal teeth from Cueva de Sidrón (Spain) and Figueira Brava (Portugal). Anthropologie 42(1): 49-52.

2.    Estebaranz F, Martínez LM, Hiraldo O, Espurz V, Bonnin A, Farrés M & Pérez-Pérez A (2004) Tooth crown size and dentine exposure in Australopithecus and early Homo: testing hypotheses of dietary related selective preasures. Anthropologie 42(1): 59-63.

3.    Galbany J & Pérez-Pérez A (2004) Buccal enamel microwear variability in Cercopithecoidea primates as a reflection of dietary habits in forested and open savanna environments. Anthropologie 42(1): 13-19.

4.    Galbany J, Martínez LM & Pérez-Pérez A (2004) Tooth replication techniques, SEM imaging and microwear analysis in Primates: methodological obstacles. Anthropologie 41(1): 5-12.

5.    Martínez LM & Pérez-Pérez A (2004) Post-mortem wear as indicator of taphonomic processes affecting enamel surfaces of hominin teeth from Laetoli and Olduvai (Tanzania): implications to dietary interpretations. Anthropologie 42(1): 37-42.

6.    Martínez LM, Galbany J & Pérez-Pérez (2004) Palaeodemography and dental microwear pattern of Homo habilis in east Africa. Anthropologie 42(1): 53-58.

7.    Pérez-Pérez A (2004) Why buccal microwear?. Anthropologie 42(1): 1-3.

8.    Pérez-Pérez A, Espurz V, Bermudez de Castro JM, de Lumley MA & Turbón D (2003) Non-occlusal dental microwear variability in a sample of middle and late pleistocene human populations from Europe and the near east. Journal of Human Evolution 44: 497-513.

9.    Romero A, Martínez Ruiz N & De Juan J (2004) Non-occlusal dental microwear in a bronze-age human sample from east Spain. Anthropologie 42(1): 65-69.

10. Ungar P, Grine F, Teaford M & Pérez-Pérez A (2001) A review of the interproximal wear grooves on fossil hominin teeth with new evidence from Olduvai Gorge. Archives of Oral Biology 46: 285-292.